In the dung beetle Onthophagus acuminatus, weapons are shown to be honest signals because

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In Onthophagus acuminatus, the presence of weapons (horns) is an honest signal because there is a trade-off in resource allocation between developing the weapons and other traits, such as eye size. This means that only individuals in good condition can afford to produce large weapons, making it an honest signal of fitness.

In the Dung Beetle Onthophagus Acuminatus, Weapons Are Shown to Be