What Evidence Led Riley and Colleagues (2005) to Suggest That

What evidence led Riley and colleagues (2005) to suggest that bees respond to the waggle dance rather than odor cues?

A) Recruits released away from the hive flew directly to the food source.
B) Recruits released away from the hive returned to it.
C) Recruits released away from the hive behaved similarly to those released at the hive.
D) All recruits flew downwind and started a zigzag search upwind for the food.
E) All recruits took much longer to fly to the food source than expected based on a straight-line route.