When Compared to Ocular Dominance Columns, How Are Orientation Columns

When compared to ocular dominance columns, how are orientation columns arranged across the cortex?

A) The center of orientation columns are found at the borders of ocular dominance columns.
B) Orientation columns do not cross the borders of ocular dominance columns.
C) The center of orientation columns are found within ocular dominance columns.
D) Orientation columns are completely separate from ocular dominance columns.
E) Orientation columns are only found in areas that have binocular input.