Mules Are Relatively Long-Lived and Hardy Organisms That Cannot, Generally

Mules are relatively long-lived and hardy organisms that cannot, generally speaking, perform successful meiosis. Consequently, which statement about mules is True?

A) They have a relative evolutionary fitness of zero.
B) Their offspring have less genetic variation than the parents.
C) Mutations cannot occur in their genomes.
D) If crossing-over happens in mules, then it must be limited to prophase of mitosis.
E) When two mules interbreed, genetic recombination cannot occur by meiotic crossing over, but only by the act of fertilization.