All of the Oxidative Steps of the Citric Acid Cycle $\alpha$

All of the oxidative steps of the citric acid cycle are linked to the reduction of NAD⁺ except the reaction catalyzed by:

A) isocitrate dehydrogenase.
B) malate dehydrogenase.
C) pyruvate dehydrogenase
D) succinate dehydrogenase.
E) the $\alpha$ -ketoglutarate dehydrogenase complex.