What Is the Difference in Function Between Simple Cells and Complex

What is the difference in function between simple cells and complex cells, in the visual cortex?

A) Simple cells respond to lines of different orientations, while complex cells respond to the whole object.
B) Simple cells are feature detectors, but complex cells are object detectors.
C) Both cells respond to orientation of lines, but simple cells can only respond to lines in specific locations.
D) Both cells are feature detectors, but simple cells can only respond to moving objects.