Which of the following is NOT a trait of a pioneer species?

Question

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Pioneer species are the first to colonize an area after a disturbance, such as a fire or landslide. They are characterized by traits such as many seeds, small seeds, and fast growth in order to quickly establish themselves in the newly open environment. However, they are not typically shade tolerant, as there is usually little to no established vegetation providing shade in a newly disturbed area.

Which of the Following Is NOT a Trait of a Pioneer