In the RNA Viruses with Segmented Genomes, Which of the Following

In the RNA viruses with segmented genomes, which of the following explains antigenic shift?

A) Lack of RNA editing by the RNA-dependent RNA polymerase.
B) Lack of proof-reading by the RNA-dependent RNA polymerase.
C) Exchange of the glycoproteins between two virions.
D) Exchange of genome segments between two virions.
E) All of the above are correct.