(Video: Ethics in animal research: The sad case of Booee the chimp) Sixteen years after leaving his trainer, it is clear that Booee has retained his ability to

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The video explicitly states that Booee is able to use sign language, as he was involved in language research and could understand and use over 300 signs. There is no mention of his ability to play the harmonica, solve math puzzles, or use complex grammar.

(Video: Ethics in Animal Research: the Sad Case of Booee