Racker and Stoeckenius used synthetic phospholipid membrane vesicles that contained a light-activated protein pump and ATP synthase to test Mitchell's chemiosmotic hypothesis. What was the conclusion of this experiment?
A) Phospholipid membrane vesicles require a proton gradient to maintain integrity.
B) ATP synthase is powered by the proton-motive force.
C) ATP synthase can generate a proton gradient.
D) Light-activated proton pumps can generate a proton gradient.
E) Light-activated proton pumps interact with ATP synthase to modulate its activity.
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