What were the most successful type of Paleozoic plants in non-swampy environments?

Name: What were the most successful type of Paleozoic plants in non-swampy environments?
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Gymnosperms, such as conifers, were able to thrive in non-swampy environments due to their ability to retain water through their needle-like leaves and deep root systems. They were also able to produce seeds, giving them a competitive advantage over other types of plants that relied on spores for reproduction.

What Were the Most Successful Type of Paleozoic Plants in Non-Swampy