Which of the following, in 1906, observed the effects of radiation by exposing rodent testicles to x-rays?

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Bergonie & Tribondeau were the first to observe the effects of radiation on rapidly dividing cells, including those of the testicles, in 1906. Ancel & Vitemberger proposed the theory of hormesis in 1943. Regaud was a French radiologist who made significant contributions to radiation therapy, but did not specifically observe the effects of radiation on testicles. Muller conducted experiments on fruit flies to demonstrate that radiation can cause mutations.

Which of the Following, in 1906, Observed the Effects of Radiation