Animals That Use Path Integration, Such as Desert Ants, Must

Animals that use path integration, such as desert ants, must collect information in order to identify the path home. What information do desert ants collect?

A) They collect directional information from the sun's position and measure distance by integrating proprioceptive information while walking.
B) They collect directional information from learned landmarks and measure distance by integrating proprioceptive information while walking.
C) They collect directional and distance information from the sun's position.
D) They collect directional information from the sun's position and distance information from learned landmarks