Which Principle Is Illustrated by Lande's Theoretical Work with Spotted

Which principle is illustrated by Lande's theoretical work with spotted owls?

A) If the availability of suitable habitat for a species goes below a critical threshold, the entire metapopulation will collapse.
B) Delay of abiotic shuffling can cause population cycles.
C) Metapopulations do not vary based on suitable habitat.
D) Introducing a small population of competitors helps keep metapopulations connected.