When Alfred Jost Removed the Ovaries from Female Rat Fetuses

When Alfred Jost removed the ovaries from female rat fetuses, the animals developed as females. However, these females were unlikely to exhibit female sexual behavior as adults because

A) typical female sexual behavior requires organizational effects of estrogens during early development.
B) typical female sexual behavior requires activational effects of estrogens during early adulthood.
C) typical female sexual behavior requires both organizational and activational effects of estrogens both during early development and in adulthood.
D) in the absence of ovaries, the brain becomes masculinized, so these animals would be expected to show male-typical sexual behavior.
E) they lacked a vagina.