To Investigate Environmental Effects on Gene Expression, Ellis and Carney

To investigate environmental effects on gene expression, Ellis and Carney (2011) did which of the following?

A) They allowed males to interact first with a male and then with a female and examined which genes were expressed during each interaction.
B) They allowed males to interact with either males or females and examined the genes that were expressed in each group.
C) They allowed females to interact with males or females and examined gene expression in each group.
D) They allowed females to interact with males and then females and examined gene expression after each interaction.
E) They allowed males and females to interact and recorded gene expression in each sex.