Current knowledge about the tempo of evolution concludes that

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Current knowledge about the tempo of evolution suggests that both gradualism and punctuated equilibrium modes of evolution have operated at different times and for different species. While some species may follow the phyletic gradualism mode, other species may follow the punctuated equilibrium mode, and at the speciation level, evolution may be almost always gradual. Therefore, the best choice is D) considering all species and long periods of time, both tempos were operating.

Current Knowledge About the Tempo of Evolution Concludes That