The main causes of species extinction include all of the following EXCEPT

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Introgression, which refers to the incorporation of genes from one species into the gene pool of another through repeated backcrossing, is not a primary cause of species extinction. Instead, it's a genetic process that can lead to gene flow between species. The other options (direct exploitation, introduced species, habitat destruction, and climate change) are well-documented causes of species extinction.

The Main Causes of Species Extinction Include All of the Following