In the golden-winged sunbird (Nectarina reichenowi), defending a territory saved the bird 780 calories a day in reduced foraging activity, but cost the bird 728 calories in defense of the territory. The net change in calories due to territorial behavior was

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The net change in calories is calculated by subtracting the cost of defending the territory (728 calories) from the calories saved by reduced foraging activity (780 calories), which equals 52 calories (780 - 728 = 52).

In the Golden-Winged Sunbird (Nectarina Reichenowi), Defending a Territory Saved