Which of the Following Statements Regarding Toma and Colleagues's Work

Which of the following statements regarding Toma and colleagues's work on the per gene and foraging in honeybees is FALSE?

A) Per gene mRNA expression was greatest in young bees that remained at the nest.
B) The per gene was used as a candidate gene involved in the control of developmental changes to foraging behavior.
C) Per gene mRNA expression was quantified in their study.
D) Young, precocious foragers had similar per mRNA levels as typical, older foragers.
E) Studies were conducted in both laboratory and natural populations of honeybees.