Hamilton's Rule States That a Gene That Codes for Assisting $\left(\sum^A_1 rb\right)+c>0$

Hamilton's rule states that a gene that codes for assisting others will increase in frequency when:

A) $\left(\sum^A_1 rb\right) +c>0$
B) $\left(\sum^A_1 rb\right) - c>0$
C) $\left(\sum^A_1 rc\right) +b>0$
D) $\left(\sum^A_1 rb\right) - c<0$
E) $\left(\sum^A_1 r\right) - bc<0$