In the discussion of cooperation via group selection, you learned that grouplevel benefits could be realized only under circumstances where there is variation in the frequency of cooperators among groups. Why must this be so? Could evolutionary forces act at the group level without such variation? Why do you suspect that group selection-based arguments have been met with such opposition? Read Wilson and Wilson's (2007) paper, "Rethinking the Theoretical Foundation of Sociobiology" (Quarterly Review of Biology, vol. 82, pp. 327-348). What is the theoretical and empirical evidence for group selection as a major evolutionary force? Describe how you might apply Wilson and Wilson's multilevel selection theory to issues such as cooperation.
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