Read the article by Muscedere and colleagues (2012) entitled "Serotonin Modulates Worker Responsiveness to Trail Pheromone in the Ant Pheidole dentate" (Journal of Comparative Physiology A, vol. 198, pp. 219-227). How did the authors determine that serotonin was an important regulator of cooperative foraging behavior in ants? Coupled with the information provided to you in the answer to Discussion Question 1 and in the Additional Readings section, generate at least two hypotheses related to how individual variation in the amount or timing of serotonin production can affect foraging success, individual fitness, and overall colony fitness in ants.
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