Read the article by Moczek & Emlen (2000) entitled "Male Horn Dimorphism in the Scarab Beetle, Onthophagus taurus: Do Alternative Reproductive Tactics Favor Alternative Phenotypes?" (Animal Behaviour, vol. 59, pp. 459-466). In this beetle species, two male morphologies exist-one with horns and one without. Horned males are incredibly adept at defending their territories, which are intricate tunnels that often contain breeding females. With that said, how can hornless male phenotypes be maintained in the population? How do you suppose the small, hornless males gain equal reproductive success as horned territorial males?
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