The research conducted by Harlow on rhesus macaques was significant because:

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Harlow's research contributed to all three of the options listed. The debate regarding nutrition versus mother-infant interaction in child development was expanded upon by Harlow's study on the importance of contact comfort in cognitive and social development. Additionally, Harlow's experiment showed that the lack of mother-infant interaction can have negative consequences on the reproductive fitness of primates. Finally, the social behavior observed in the rhesus macaques by Harlow can be extrapolated to better understand human behaviors and relationships.

The Research Conducted by Harlow on Rhesus Macaques Was Significant