What respiratory adaptation was important for the invasion of land by arthropods?

Question

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Arthropods were able to invade land thanks to the development of body systems that allowed them to avoid desiccation. One of the key respiratory adaptations that played a role in this was the ability to close spiracles, which are openings in their exoskeleton that allow for gas exchange. By closing the spiracles, arthropods can reduce water loss in their dry terrestrial environments.

What Respiratory Adaptation Was Important for the Invasion of Land