Molecules Called Uncouplers Allow Protons to Cross the Inner Mitochondrial

Molecules called uncouplers allow protons to cross the inner mitochondrial membrane.What effect would this have on ATP production in the mitochondria?

A) Reduction, because electrons would not be able to move through the electron transport chain
B) Reduction, because protons would not be available to pass through the ATPase
C) No effect, because protons cross the outer mitochondrial membrane to produce ATP
D) Increase, because this would spontaneously establish a proton gradient
E) Increase, because this would make more oxygen available in the mitochondrial matrix