What Hypothesis Were Ellison and Gotelli Testing When They Manipulated

What hypothesis were Ellison and Gotelli testing when they manipulated concentrations of nutrients available to pitcher plants?

A) Providing extra light would increase leaf production.
B) Providing extra nutrients would increase photosynthetic leaf surface area.
C) Providing extra light would increase the need for nutrients.
D) Providing extra nutrients would reduce the need for extra leaves.
E) Cutting leaves would diminish the need for more nutrients.