Electron Transport Within NADH-Q Reductase, Cytochrome Reductase, and Cytochrome Oxidase

Electron transport within NADH-Q reductase, cytochrome reductase, and cytochrome oxidase can be coupled to proton transport from the mitochondrial matrix to the space between the inner and outer mitochondrial membranes.This is because those protein complexes

A) are in the mitochondrial matrix.
B) span the inner mitochondrial membrane.
C) are in the space between the inner and outer mitochondrial membranes.
D) remain in the cytoplasm.
E) loosely attach to the inner mitochondrial membrane.