Which Structures Characteristic of Vascular Plants Developed First in the Lycophytes

Which structures characteristic of vascular plants developed first in the lycophytes?

A) A thick, waxy cuticle to prevent desiccation and pores called stomata to allow for gas exchange
B) Megaphylls and a specialized arrangement of vascular tissue to allow differentiation of water and sugar transport
C) An independent sporophyte and true roots that branch dichotomously
D) A protected embryo and pores called stomata to allow for gas exchange
E) True roots that branch dichotomously and a persistently green sporophyte