During the Process of Succession, Early Species Tend to Be

During the process of succession, early species tend to be facilitators, while many later species compete with each other and act more as inhibitors.Yet, species diversity tends to increase as succession continues.How is it possible for high species diversity to coexist with inhibitory interactions in mature communities?

A) Mature communities can maintain high diversity by adding large, perennial species that are adapted to overcome competition.
B) Mature communities have a combination of both facilitating and inhibiting interactions, which act together to maintain diversity.
C) Facilitation is only required in young communities just beginning succession; mature communities are designed to live with only inhibitory interactions.
D) Both facilitation and inhibition are present in equal amounts throughout succession, but inhibition is harder to detect in young communities.
E) Later successional species are much more stress-tolerant than earlier species and can therefore withstand inhibitory interactions.