In Harry Harlow's early work with infant monkeys, he noticed that when they were raised with adequate food but without any attention from or contact with their mothers or other adult monkeys, they usually:

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Harry Harlow (1959) noted that when monkeys in his lab were raised without their mothers, they often developed maladapted behaviors. They were easily frightened, irritable, and reluctant to eat or play, despite the fact that they had a nutritious diet and were otherwise well cared for. Some monkeys even died. Obviously, these baby monkeys needed something more than regular feeding to thrive and develop.

In Harry Harlow's Early Work with Infant Monkeys, He Noticed