What is Darwin's "measure of success" in Origin of Species?

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In Origin of Species, Darwin's "measure of success" is the ability to reproduce fertile offspring. This is because the passing on of genes to the next generation through successful reproduction is crucial for the evolution of a population. Other choices such as spiritual superiority, economic affluence, and natural evolution are not explicitly mentioned as Darwin's measure of success, and while struggle for survival is important for natural selection, it is not the measure of success itself but rather a driving force behind it.

What Is Darwin's "Measure of Success" in Origin of Species