Which of the Following Statements Is FALSE

Which of the following statements is FALSE?

A) All of the protein complexes of the mitochondrial respiratory chain are bound in the inner mitochondrial membrane.
B) Cytochrome c and coenzyme Q are both soluble electron carriers that are loosely attached to the outside of the inner mitochondrial membrane.
C) Oxygen is reduced to water at complex IV of the mitochondrial respiratory chain.
D) The citric acid cycle is linked directly to the mitochondrial respiratory chain at the site of complex II.
E) During electron transport protons are pumped across the inner mitochondrial membrane from the matrix to the intermembrane space.