Stacked and unstacked regions of the thylakoid are arranged such that:

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In chloroplasts, the thylakoid membrane is differentiated into stacked (grana) and unstacked (stroma lamellae) regions. Photosystem I is primarily found in the unstacked regions, while Photosystem II is located in the stacked regions. ATP synthase is also mainly found in the unstacked regions, facilitating ATP synthesis using the proton gradient established across the thylakoid membrane.

Stacked and Unstacked Regions of the Thylakoid Are Arranged Such