What would happen if complexes I-IV of the electron transport chain pumped protons in the opposite direction?

Question

Quizplus · Accepted Answer

The pumping of protons in the opposite direction by complexes I-IV of the electron transport chain would result in a decrease in the proton gradient across the inner mitochondrial membrane. This gradient is necessary for the synthesis of ATP by ATP synthase. Without a sufficient proton gradient, no ATP would be synthesized.

What Would Happen If Complexes I-IV of the Electron Transport