Male Northern Elephant Seals (Mirounga Angustirostris) Compete with One Another

Male northern elephant seals (Mirounga angustirostris) compete with one another for territories along the beach. Males defend these territories throughout the breeding season. The males that win the male-male competitions defend the "best" territories, and have access to females that stay in that territory of the beach. The females land on the beach during mating season, choosing which territory of the beach to join. They then disproportionately breed with the male that defends that area of the beach. At what point may intersexual selection occur in these elephant seals?

A) when the female is swimming to the beach
B) when the female lands on the beach and chooses her resting location
C) when the male defends his group of females against a different male
D) when the male mates with a female within his group
E) when the female leaves the beach after mating with a male