Which of the Following Statements About the Chemiosmotic Theory Is

Which of the following statements about the chemiosmotic theory is false?

A) Electron transfer in mitochondria is accompanied by an asymmetric release of protons on one side of the inner mitochondrial membrane.
B) Energy is conserved as a transmembrane pH gradient.
C) Oxidative phosphorylation cannot occur in membrane-free preparations.
D) The effect of uncoupling reagents is a consequence of their ability to carry protons through membranes.
E) The membrane ATPase,which plays an important role in other hypotheses for energy coupling,has no significant role in the chemiosmotic theory.