Joseph Connell discovered that Balanus barnacles were excluded from the upper intertidal zone by ________, while Chthalamus barnacles were excluded from the middle intertidal zone by ________.
A) competition from mussels; prolonged immersion in water
B) competition from mussels; competition with Balanus
C) prolonged exposure to air; competition with Balanus
D) prolonged exposure to air; competition with algae
E) predation by wading birds; predation by starfish
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