Which Conclusion Was Not Made from the Seminal 1979 Paper

Which conclusion was not made from the seminal 1979 paper by Gould and Lewontin?

A) Natural selection in the present environment is just one of several processes by which a species may come to exhibit a trait.
B) When physiologists refer to a trait as an adaptation, they are making a hypothesis that natural selection has occurred.
C) Data must be gathered in order to assess whether adaptation is likely to have occurred.
D) Indirect evidence cannot be used to support the hypothesis of adaptation.