What Prediction of the Optimal Patch Model Did Tome (1988)

What prediction of the optimal patch model did Tome (1988) test on ruddy ducks?

A) As travel time increases, individuals should spend more time in and harvest more items from a patch
B) Quitting harvest rates will be higher in rich compared to poor patches
C) Giving-up densities should vary with food patch quality
D) The marginal benefit of feeding should equal the missed opportunity costs of exploiting a patch
E) The cumulative gain curve should get steeper as more time is spent in a patch