What Prediction Did Biernaske, Walker, and Gegear (2009) Test to Investigate

What prediction did Biernaske, Walker, and Gegear (2009) test to investigate Bayesian estimation in foraging bees?

A) Bees trained on different environments will have different marginal benefits of feeding
B) Bees trained on different environments will exhibit different GUDs
C) Variation in behavior will match environmental variation
D) Bees trained in a high-variance environment will exhibit high variation in behavior
E) Bees trained on different environments will exhibit different patch-leaving behaviors