Two species of plants that diverged 25 million years ago have accumulated an estimated 18 nucleotide differences in a gene.Species A has an average population size of 2 million, while species B has an average population size of 0.8 million.Both of these plants are annuals, with one generation per year.Assuming that the substitutions are all neutral, the estimated neutral mutation rate for this gene is _______ per million generations.
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