Use the Following Abstract from Theoretical Population Biology to Answer

Use the following abstract from Theoretical Population Biology to answer the question.
Abstract:
We derive measures for assessing the value of an individual habitat fragment for the dynamics and persistence of a metapopulation living in a network of many fragments. We demonstrate that the most appropriate measure of fragment value depends on the question asked. Specifically, we analyze four alternative measures: the contribution of a fragment to the metapopulation capacity of the network, to the equilibrium metapopulation size, to the expected time to metapopulation extinction and the long-term contribution of a fragment to colonization events in the network. The latter measure is comparable to density-dependent measures in general matrix population theory, though some differences are introduced by the fact that "density dependence" is spatially localized in the metapopulation context. We show that the value of a fragment depends not only on the properties of the landscape but also on the properties of the species. Most importantly, variation in fragment values between the habitat fragments is greatest in the case of rare species that occur close to the extinction threshold, as these species are likely to be restricted to the most favorable parts of the landscape. We expect that the measures of habitat fragment described and analyzed here have applications in landscape ecology and in conservation biology.
Copyright © 2003 Elsevier Inc. All rights reserved. (Otso Ovaskainen and Ilkka Hanski. 2003. How much does an individual habitat fragment contribute to metapopulation dynamics and persistence? Theoretical Population Biology 64:481-95.)
One measure for the value of the patch was given by the long-term contribution of a fragment to colonization events in the network. How do the properties of a landscape and the properties of a species affect the value of a patch? The value of the fragment depends ________.

A) on the properties of the landscape and the properties of the species
B) only on the properties of the landscape and not on the properties of the species
C) not on the properties of the landscape but only on the properties of the species
D) on neither the properties of the landscape nor on the properties of the species